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Territoriality and Inter-Pack Aggression in Gray Wolves: Shaping a Social Carnivore’s Life History

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Kira A. Cassidy, Douglas W. Smith, L. David Mech, Daniel R. MacNulty, Daniel R. Stahler and Matthew C. Metz (2016). Territoriality and Inter-Pack Aggression in Gray Wolves: Shaping a Social Carnivore's Life History. Yellowstone Science 24(1): 37–42.

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  • Wolf-on-wolf conflict accounted for about two-thirds of known natural mortality among Yellowstone National Park collared wolves (1995–2015; Figure 1).
  • The authors report that, in the observed Yellowstone inter-pack conflicts, the presence of older wolves was a better indicator of success than total pack size.


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