Wolf Research

Research publications by L. David Mech. Related reference: Mechโ€™s explanation of โ€œalphaโ€ terminology.

Dominance Relationships in a Family Pack of Captive Arctic Wolves (Canis lupus arctos): The Influence of Competition for Food, Age and Sex

Publication Simona Cafazzo, Martina Lazzaroni and Sarah Marshall-Pescini (2016). Dominance Relationships in a Family Pack of Captive Arctic Wolves (Canis lupus arctos): The Influence of Competition for Food, Age and Sex. PeerJ 4: e2707. Original sourceโ€ฆ

In Wolves, Play Behaviour Reflects the Partners’ Affiliative and Dominance Relationship

Publication Simona Cafazzo, Sarah Marshall-Pescini, Jennifer L. Essler, Zsรณfia Virรกnyi, Kurt Kotrschal and Friederike Range (2018). In Wolves, Play Behaviour Reflects the Partners' Affiliative and Dominance Relationship. Animal Behaviour 141: 137โ€“150. Original source (DOI) ยท Downloadโ€ฆ

Territoriality and Inter-Pack Aggression in Gray Wolves: Shaping a Social Carnivore’s Life History

Publication Kira A. Cassidy, Douglas W. Smith, L. David Mech, Daniel R. MacNulty, Daniel R. Stahler and Matthew C. Metz (2016). Territoriality and Inter-Pack Aggression in Gray Wolves: Shaping a Social Carnivore's Life History. Yellowstone Scienceโ€ฆ

Play Behavior in Wolves: Using the โ€˜50:50โ€™ Rule to Test for Egalitarian Play Styles

Source Jennifer L. Essler, Simona Cafazzo, Sarah Marshall-Pescini, Zsรณfia Virรกnyi, Kurt Kotrschal, and Friederike Range (2016). Play Behavior in Wolves: Using the โ€˜50:50โ€™ Rule to Test for Egalitarian Play Styles. PLOS ONE. 11(5), e0154150. Research article.โ€ฆ

The Power of Discourse: Associations between Trainersโ€™ Speech and the Responses of Socialized Wolves and Dogs to Training

Publication Melissa Gabriela Bravo Fonseca et al. The Power of Discourse: Associations between Trainersโ€™ Speech and the Responses of Socialized Wolves and Dogs to Training. Animals 13:1071 (2023). [11] DOI: 10.3390/ani13061071 Melissa Gabriela Bravo Fonseca,Heron Oliveiraโ€ฆ

Differences in post-conflict behaviour in wolves and dog-admixed wolves

Authors: Federica Amici, Katja Liebal, Linda S. Oรฑa, Arne Gretschzel, Tina Altdรถrfer, Mattia Iacuzzi, George Kamanga, Manon Delaunay, Paolo Ciucci. Publication: Royal Society Open Science, 13, 260767 (2026). Published online 21 October 2026. Original study: DOI:โ€ฆ